Southern operates a train from Lewes to Firle hourly. Tickets cost $2 - $3 and the journey takes 5 min. Alternatively, Seaford & District operates a bus from Lewes to Firle 3 times a day. Tickets cost $4 and the journey takes 15 min. Cuckmere Buses also services this route once daily.
From Lewes train to Glynde. 5 min £1. From Lewes, Bus Station bus to Firle, Stores. 15 min £2. From Lewes taxi to Firle. 7 min £13. From Lewes Uber to Firle. 7 min £17. From Lewes walk to Firle. 1 h 29 min.